By Richard D Smallwood

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Folia Primatol 66:126–136. Kram R, Taylor CR (1990) Energetics of running: a new perspective. Nature 346:265–267. Larney E, Larson SG (2004) Limb compliance during walking: comparisons of elbow and knee yield across quadrupedal primates and in other mammals. Am J Phys Anthropol 125:42–50. Larson SG (1998) Unique aspects of quadrupedal locomotion in nonhuman primates. In: Strasser E, Fleagle JG, Rosenberger AL, McHenry HM (eds), Primate Locomotion: Recent Advances. Plenum, New York, pp 157–173. Larson SG, Schmitt D, Lemelin P, Hamrick MW (1999) The uniqueness of primate forelimb posture during quadrupedal locomotion.

This chapter focuses on two ways in which stability can be regulated: 1) by choosing gaits or timing of footfalls that maintain as many limbs as possible in contact with the substrate (Cartmill et al. 2002) and 2) avoiding any whole-body aerial phase (Schmitt et al. 2006). Peak Ground Forces Avoiding catastrophic limb fracture as well as smaller microfractures and joint damage is another critical performance variable. It is well known that by combining limb biomechanical properties and limb posture, animals consistently produce safety factors that allow resistance to fracture two to four times higher than needed for normal locomotion (Biewener 1989, 1990).

J Hum Evol 46:237– 252. Schmitt D, Larson SG, Stern JT (1994) Serratus ventralis function in vervet monkeys: Are primate quadrupeds unique? J Zool 232:215–230. Schmitt D, Lemelin P (2002) The origins of primate locomotion: gait mechanics of the woolly opossum. Am J Phys Anthropol 118:231–238. 2 Laboratory and Field Biomechanics 27 Schmitt D, Lemelin P (2004) Locomotor mechanics of the slender loris (Loris tardigradus). J Hum Evol 47:85–94. Schmitt D, Cartmill M, Hanna J, Griffin T, Lemelin P (2006) Adaptive value of ambling gait in primates and other mammals.

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